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id int(11) 475 Column Stats
subject varchar(80) 14 Column Stats
title varchar(250) 139 Column Stats
ordinal int(11) 30 Column Stats
description text 474 Column Stats

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06 - Difficutiles in Theory 06-13 - Summary: The Law of Unity of Type and of the Conditions of Existence Embraced by the Theory of Natural Selection 40 Although the belief that an organ so perfect as the eye could have been formed by natural selection, is enough to stagger any one; yet in the case of any organ, if we know of a long series of gradations in complexity, each good for its possessor, then, under changing conditions of life, there is no logical impossibility in the acquirement of any conceivable degree of perfection through natural selection.
eye
eye


In the cases in which we know of no intermediate or transitional states, we should be extremely cautious in concluding that none can have existed, for the metamorphoses of many organs show what wonderful changes in function are at least possible.

For instance, a swimbladder has apparently been converted into an air-breathing lung.
Swim Bladder
Swim Bladder

lungs
lungs


The same organ having performed simultaneously very different functions, and then having been in part or in whole specialised for one function; and two distinct organs having performed at the same time the same function, the one having been perfected whilst aided by the other, must often have largely facilitated transitions.
14 - Recapitulation and Conclusion 14-03 - Causes of the general belief in the immutability of species 130 Although I am fully convinced of the truth of the views given in this volume under the form of an abstract, I by no means expect to convince experienced naturalists whose minds are stocked with a multitude of facts all viewed, during a long course of years, from a point of view directly opposite to mine.

It is so easy to hide our ignorance under such expressions as the `plan of creation,' `unity of design,' &c., and to think that we give an explanation when we only restate a fact.

Any one whose disposition leads him to attach more weight to unexplained difficulties than to the explanation of a certain number of facts will certainly reject my theory.

A few naturalists, endowed with much flexibility of mind, and who have already begun to doubt on the immutability of species, may be influenced by this volume; but I look with confidence to the future, to young and rising naturalists, who will be able to view both sides of the question with impartiality.

Whoever is led to believe that species are mutable will do good service by conscientiously expressing his conviction; for only thus can the load of prejudice by which this subject is overwhelmed be removed.

Several eminent naturalists have of late published their belief that a multitude of reputed species in each genus are not real species; but that other species are real, that is, have been independently created.

This seems to me a strange conclusion to arrive at.

They admit that a multitude of forms, which till lately they themselves thought were special creations, and which are still thus looked at by the majority of naturalists, and which consequently have every external characteristic feature of true species, -- they admit that these have been produced by variation, but they refuse to extend the same view to other and very slightly different forms.

Nevertheless they do not pretend that they can define, or even conjecture, which are the created forms of life, and which are those produced by secondary laws.

They admit variation as a vera causa in one case, they arbitrarily reject it in another, without assigning any distinction in the two cases.

The day will come when this will be given as a curious illustration of the blindness of preconceived opinion.

These authors seem no more startled at a miraculous act of creation than at an ordinary birth.

But do they really believe that at innumerable periods in the earth's history certain elemental atoms have been commanded suddenly to flash into living tissues?

Do they believe that at each supposed act of creation one individual or many were produced?

Were all the infinitely numerous kinds of animals and plants created as eggs or seed, or as full grown?

egg
egg

seeds
seeds


and in the case of mammals, were they created bearing the false marks of nourishment from the mother's womb?

mammals
mammals


Although naturalists very properly demand a full explanation of every difficulty from those who believe in the mutability of species, on their own side they ignore the whole subject of the first appearance of species in what they consider reverent silence.
02 - Variations Under Nature 02-04 - Wide-ranging, much diffused, and common Species vary most 20 Alphonse de Candolle and others have shown that plants which have very wide ranges generally present varieties; and this might have been expected, as they are exposed to diverse physical conditions, and as they come into competition (which, as we shall hereafter see, is an equally or more important circumstance) with different sets of organic beings.

Alphonse de Candolle
Alphonse de Candolle


But my tables further show that, in any limited country, the species which are the most common, that is abound most in individuals, and the species which are most widely diffused within their own country (and this is a different consideration from wide range, and to a certain extent from commonness), oftenest give rise to varieties sufficiently well marked to have been recorded in botanical works.

Hence it is the most flourishing, or, as they may be called, the dominant species,- those which range widely, are the most diffused in their own country, and are the most numerous in individuals,- which oftenest produce well-marked varieties, or, as I consider them, incipient species.

And this, perhaps, might have been anticipated; for as varieties, in order to become in any degree permanent, necessarily have to struggle with the other inhabitants of the country, the species which are already dominant will be the most likely to yield offspring, which, though in some slight degree modified, still inherit those advantages that enabled their parents to become dominant over their compatriots.

In these remarks on predominance, it should be understood that reference is made only to the forms which come into competition with each other, and more especially to the members of the same genus or class having nearly similar habits of life.
12 - Geographical Distribution -- continued 12-40 - On the relations of the inhabitants of islands to those of the nearest mainland 30 All the foregoing remarks on the inhabitants of oceanic islands, namely, the scarcity of kinds -- the richness in endemic forms in particular classes or sections of classes, the absence of whole groups, as of batrachians, and of terrestrial mammals notwithstanding the presence of aerial bats, the singular proportions of certain orders of plants, herbaceous forms having been developed into trees, &c., seem to me to accord better with the view of occasional means of transport having been largely efficient in the long course of time, than with the view of all our oceanic islands having been formerly connected by continuous land with the nearest continent; for on this latter view the migration would probably have been more complete; and if modification be admitted, all the forms of life would have been more equally modified, in accordance with the paramount importance of the relation of organism to organism.

I do not deny that there are many and grave difficulties in understanding how several of the inhabitants of the more remote islands, whether still retaining the same specific form or modified since their arrival, could have reached their present homes.

But the probability of many islands having existed as halting-places, of which not a wreck now remains, must not be overlooked.

I will here give a single instance of one of the cases of difficulty.

Almost all oceanic islands, even the most isolated and smallest, are inhabited by land-shells, generally by endemic species, but sometimes by species found elsewhere.

Land Shell
Land Shell


Dr. Aug. A. Gould has given several interesting cases in regard to the land-shells of the islands of the Pacific.

Augustus Addison Gould
Augustus Addison Gould

Pacific Ocean
Pacific Ocean


Now it is notorious that land-shells are very easily killed by salt; their eggs, at least such as I have tried, sink in sea-water and are killed by it.

Yet there must be, on my view, some unknown, but highly efficient means for their transportal.

Would the just-hatched young occasionally crawl on and adhere to the feet of birds roosting on the ground, and thus get transported? It occurred to me that land-shells, when hybernating and having a membranous diaphragm over the mouth of the shell, might be floated in chinks of drifted timber across moderately wide arms of the sea.

And I found that several species did in this state withstand uninjured an immersion in sea-water during seven days: one of these shells was the Helix pomatia, and after it had again hybernated I put it in sea-water for twenty days, and it perfectly recovered.

Helix Pomatia
Helix Pomatia


As this species has a thick calcareous operculum, I removed it, and when it had formed a new membranous one, I immersed it for fourteen days in sea-water, and it recovered and crawled away: but more experiments are wanted on this head.