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|OriginOfSpecies - 475 Rows|
|13 - Mutual Affinities of Organic Beings: Morphology: Embryology: Rudimentary Or||13-03 - Rules and difficulties in classification, explained on the theory of descent with modification||40||
Any one of these characters singly is frequently of more than generic importance, though here even when all taken together they appear insufficient to separate Cnestis from Connarus.' To give an example amongst insects, in one great division of the Hymenoptera, the antennae, as Westwood has remarked, are most constant in structure; in another division they differ much, and the differences are of quite subordinate value in classification; yet no one probably will say that the antennae in these two divisions of the same order are of unequal physiological importance.
Any number of instances could be given of the varying importance for classification of the same important organ within the same group of beings.
Again, no one will say that rudimentary or atrophied organs are of high physiological or vital importance; yet, undoubtedly, organs in this condition are often of high value in classification.
No one will dispute that the rudimentary teeth in the upper jaws of young ruminants, and certain rudimentary bones of the leg, are highly serviceable in exhibiting the close affinity between Ruminants and Pachyderms.
Robert Brown has strongly insisted on the fact that the rudimentary florets are of the highest importance in the classification of the Grasses.
|14 - Recapitulation and Conclusion||14-05 - Effects of its adoption on the study of Natural history||40||
Embryology will reveal to us the structure, in some degree obscured, of the prototypes of each great class.
When we can feel assured that all the individuals of the same species, and all the closely allied species of most genera, have within a not very remote period descended from one parent, and have migrated from some one birthplace; and when we better know the many means of migration, then, by the light which geology now throws, and will continue to throw, on former changes of climate and of the level of the land, we shall surely be enabled to trace in an admirable manner the former migrations of the inhabitants of the whole world.
Even at present, by comparing the differences of the inhabitants of the sea on the opposite sides of a continent, and the nature of the various inhabitants of that continent in relation to their apparent means of immigration, some light can be thrown on ancient geography.
The noble science of Geology loses glory from the extreme imperfection of the record.
The crust of the earth with its embedded remains must not be looked at as a well-filled museum, but as a poor collection made at hazard and at rare intervals.
The accumulation of each great fossiliferous formation will be recognised as having depended on an unusual concurrence of circumstances, and the blank intervals between the successive stages as having been of vast duration.
But we shall be able to gauge with some security the duration of these intervals by a comparison of the preceding and succeeding organic forms.
We must be cautious in attempting to correlate as strictly contemporaneous two formations, which include few identical species, by the general succession of their forms of life.
|07 - Instinct||07-04 - Instincts variable||40||
As some degree of variation in instincts under a state of nature, and the inheritance of such variations, are indispensable for the action of natural selection, as many instances as possible ought to have been here given; but want of space prevents me.
I can only assert, that instincts certainly do vary for instance, the migratory instinct, both in extent and direction, and in its total loss.
So it is with the nests of birds, which vary partly in dependence on the situations chosen, and on the nature and temperature of the country inhabited, but often from causes wholly unknown to us: Audubon has given several remarkable cases of differences in nests of the same species in the northern and southern United States.
Fear of any particular enemy is certainly an instinctive quality, as may be seen in nestling birds, though it is strengthened by experience, and by the sight of fear of the same enemy in other animals.
But fear of man is slowly acquired, as I have elsewhere shown, by various animals inhabiting desert islands; and we may see an instance of this, even in England, in the greater wildness of all our large birds than of our small birds; for the large birds have been most persecuted by man.
We may safely attribute the greater wildness of our large birds to this cause; for in uninhabited islands large birds are not more fearful than small; and the magpie, so wary in England, is tame in Norway, as is the hooded crow in Egypt.
That the general disposition of individuals of the same species, born in a state of nature, is extremely diversified, can be shown by a multitude of facts.
Several cases also, could be given, of occasional and strange habits in certain species, which might, if advantageous to the species, give rise, through natural selection, to quite new instincts.
But I am well aware that these general statements, without facts given in detail, can produce but a feeble effect on the reader's mind. I can only repeat my assurance, that I do not speak without good evidence.
|14 - Recapitulation and Conclusion||14-02 - Recapitulation of the general and special circumstances in its favour||40||
If then we have under nature variability and a powerful agent always ready to act and select, why should we doubt that variations in any way useful to beings, under their excessively complex relations of life, would be preserved, accumulated, and inherited?
Why, if man can by patience select variations most useful to himself, should nature fail in selecting variations useful, under changing conditions of life, to her living products?
What limit can be put to this power, acting during long ages and rigidly scrutinising the whole constitution, structure, and habits of each creature, -- favouring the good and rejecting the bad?
I can see no limit to this power, in slowly and beautifully adapting each form to the most complex relations of life.
The theory of natural selection, even if we looked no further than this, seems to me to be in itself probable.