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07 - Instinct 07-11 - Summary 10 Summary.

I have endeavoured briefly in this chapter to show that the mental qualities of our domestic animals vary, and that the variations are inherited.

Still more briefly I have attempted to show that instincts vary slightly in a state of nature.

No one will dispute that instincts are of the highest importance to each animal.

Therefore I can see no difficulty, under changing conditions of life, in natural selection accumulating slight modifications of instinct to any extent, in any useful direction. In some cases habit or use and disuse have probably come into play.

I do not pretend that the facts given in this chapter strengthen in any great degree my theory; but none of the cases of difficulty, to the best of my judgment, annihilate it.

On the other hand, the fact that instincts are not always absolutely perfect and are liable to mistakes; that no instinct has been produced for the exclusive good of other animals, but that each animal takes advantage of the instincts of others; that the canon in natural history, of 'natura non facit saltum' is applicable to instincts as well as to corporeal structure, and is plainly explicable on the foregoing views, but is otherwise inexplicable, all tend to corroborate the theory of natural selection.

This theory is, also, strengthened by some few other facts in regard to instincts; as by that common case of closely allied, but certainly distinct, species, when inhabiting distant parts of the world and living under considerably different conditions of life, yet often retaining nearly the same instincts.

For instance, we can understand on the principle of inheritance, how it is that the thrush of South America lines its nest with mud, in the same peculiar manner as does our British thrush: how it is that the male wrens (Troglodytes) of North America, build 'cock-nests,' to roost in, like the males of our distinct Kitty-wrens, a habit wholly unlike that of any other known bird.

thrush
thrush

wren
wren


Finally, it may not be a logical deduction, but to my imagination it is far more satisfactory to look at such instincts as the young cuckoo ejecting its foster-brothers, ants making slaves, -- the larvae of ichneumonidae feeding within the live bodies of caterpillars, not as specially endowed or created instincts, but as small consequences of one general law, leading to the advancement of all organic beings, namely, multiply, vary, let the strongest live and the weakest die.

cuckoo
cuckoo

ichneumonidae
ichneumonidae

caterpillar
caterpillar
08 - Hybridism 08-09 - Summary 10 Summary of Chapter.

First crosses between forms sufficiently distinct to be ranked as species, and their hybrids, are very generally, but not universally, sterile.

The sterility is of all degrees, and is often so slight that the two most careful experimentalists who have ever lived, have come to diametrically opposite conclusions in ranking forms by this test.

The sterility is innately variable in individuals of the same species, and is eminently susceptible of favourable and unfavourable conditions.

The degree of sterility does not strictly follow systematic affinity, but is governed by several curious and complex laws.

It is generally different, and sometimes widely different, in reciprocal crosses between the same two species.

It is not always equal in degree in a first cross and in the hybrid produced from this cross.

In the same manner as in grafting trees, the capacity of one species or variety to take on another, is incidental on generally unknown differences in their vegetative systems, so in crossing, the greater or less facility of one species to unite with another, is incidental on unknown differences in their reproductive systems.

graft
graft

tree
tree


There is no more reason to think that species have been specially endowed with various degrees of sterility to prevent them crossing and blending in nature, than to think that trees have been specially endowed with various and somewhat analogous degrees of difficulty in being grafted together in order to prevent them becoming inarched in our forests.

The sterility of first crosses between pure species, which have their reproductive systems perfect, seems to depend on several circumstances; in some cases largely on the early death of the embryo.

embryo
embryo


The sterility of hybrids, which have their reproductive systems imperfect, and which have had this system and their whole organisation disturbed by being compounded of two distinct species, seems closely allied to that sterility which so frequently affects pure species, when their natural conditions of life have been disturbed.

This view is supported by a parallelism of another kind; namely, that the crossing of forms only slightly different is favourable to the vigour and fertility of their offspring; and that slight changes in the conditions of life are apparently favourable to the vigour and fertility of all organic beings.

It is not surprising that the degree of difficulty in uniting two species, and the degree of sterility of their hybrid-offspring should generally correspond, though due to distinct causes; for both depend on the amount of difference of some kind between the species which are crossed.

Nor is it surprising that the facility of effecting a first cross, the fertility of the hybrids produced, and the capacity of being grafted together though this latter capacity evidently depends on widely different circumstances should all run, to a certain extent, parallel with the systematic affinity of the forms which are subjected to experiment; for systematic affinity attempts to express all kinds of resemblance between all species.

First crosses between forms known to be varieties, or sufficiently alike to be considered as varieties, and their mongrel offspring, are very generally, but not quite universally, fertile.

Nor is this nearly general and perfect fertility surprising, when we remember how liable we are to argue in a circle with respect to varieties in a state of nature; and when we remember that the greater number of varieties have been produced under domestication by the selection of mere external differences, and not of differences in the reproductive system.

In all other respects, excluding fertility, there is a close general resemblance between hybrids and mongrels.

Finally, then, the facts briefly given in this chapter do not seem to me opposed to, but even rather to support the view, that there is no fundamental distinction between species and varieties.

jaglion
jaglion

liger
liger

zeedonk
zeedonk
10 - On The Geological Succession of Organic Beings 10-02 - On their different rates of change 10 Species of different genera and classes have not changed at the same rate, or in the same degree.

In the oldest tertiary beds a few living shells may still be found in the midst of a multitude of extinct forms.

Sea Shell
Sea Shell


Falconer has given a striking instance of a similar fact, in an existing crocodile associated with many strange and lost mammals and reptiles in the sub-Himalayan deposits.

Hugh Falconer
Hugh Falconer

crocodile
crocodile

mammals
mammals

Reptile
Reptile

Himalaya
Himalaya


The Silurian Lingula differs but little from the living species of this genus; whereas most of the other Silurian Molluscs and all the Crustaceans have changed greatly.

lingula
lingula

mollusca
mollusca

crustacean
crustacean


The productions of the land seem to change at a quicker rate than those of the sea, of which a striking instance has lately been observed in Switzerland.

Switzerland
Switzerland


There is some reason to believe that organisms, considered high in the scale of nature, change more quickly than those that are low: though there are exceptions to this rule.

The amount of organic change, as Pictet has remarked, does not strictly correspond with the succession of our geological formations; so that between each two consecutive formations, the forms of life have seldom changed in exactly the same degree.

Yet if we compare any but the most closely related formations, all the species will be found to have undergone some change.

When a species has once disappeared from the face of the earth, we have reason to believe that the same identical form never reappears.

The strongest apparent exception to this latter rule, is that of the so-called `colonies' of M. Barrande, which intrude for a period in the midst of an older formation, and then allow the pre-existing fauna to reappear; but Lyell's explanation, namely, that it is a case of temporary migration from a distinct geographical province, seems to me satisfactory.

Joachim Barrande
Joachim Barrande

Sir Charles Lyell
Sir Charles Lyell


These several facts accord well with my theory.

I believe in no fixed law of development, causing all the inhabitants of a country to change abruptly, or simultaneously, or to an equal degree.

The process of modification must be extremely slow.

The variability of each species is quite independent of that of all others.

Whether such variability be taken advantage of by natural selection, and whether the variations be accumulated to a greater or lesser amount, thus causing a greater or lesser amount of modification in the varying species, depends on many complex contingencies, on the variability being of a beneficial nature, on the power of intercrossing, on the rate of breeding, on the slowly changing physical conditions of the country, and more especially on the nature of the other inhabitants with which the varying species comes into competition.

Hence it is by no means surprising that one species should retain the same identical form much longer than others; or, if changing, that it should change less.

We see the same fact in geographical distribution; for instance, in the land-shells and coleopterous insects of Madeira having come to differ considerably from their nearest allies on the continent of Europe, whereas the marine shells and birds have remained unaltered.

Land Shell
Land Shell

Madeira
Madeira

europe
europe

Sea Shell
Sea Shell

bird
bird


We can perhaps understand the apparently quicker rate of change in terrestrial and in more highly organised productions compared with marine and lower productions, by the more complex relations of the higher beings to their organic and inorganic conditions of life, as explained in a former chapter.

When many of the inhabitants of a country have become modified and improved, we can understand, on the principle of competition, and on that of the many all-important relations of organism to organism, that any form which does not become in some degree modified and improved, will be liable to be exterminated.

Hence we can see why all the species in the same region do at last, if we look to wide enough intervals of time, become modified; for those which do not change will become extinct.

In members of the same class the average amount of change, during long and equal periods of time, may, perhaps, be nearly the same; but as the accumulation of long-enduring fossiliferous formations depends on great masses of sediment having been deposited on areas whilst subsiding, our formations have been almost necessarily accumulated at wide and irregularly intermittent intervals; consequently the amount of organic change exhibited by the fossils embedded in consecutive formations is not equal.

Each formation, on this view, does not mark a new and complete act of creation, but only an occasional scene, taken almost at hazard, in a slowly changing drama.
09 - On the Imperfection of the Geological Record 09-02 - On the nature of extinct intermediate varieties; on their number 10 So with natural species, if we look to forms very distinct, for instance to the horse and tapir, we have no reason to suppose that links ever existed directly intermediate between them, but between each and an unknown common parent.

horse
horse

tapir
tapir


The common parent will have had in its whole organisation much general resemblance to the tapir and to the horse; but in some points of structure may have differed considerably from both, even perhaps more than they differ from each other.

Hence in all such cases, we should be unable to recognise the parent-form of any two or more species, even if we closely compared the structure of the parent with that of its modified descendants, unless at the same time we had a nearly perfect chain of the intermediate links.

It is just possible by my theory, that one of two living forms might have descended from the other; for instance, a horse from a tapir; and in this case direct intermediate links will have existed between them.

But such a case would imply that one form had remained for a very long period unaltered, whilst its descendants had undergone a vast amount of change; and the principle of competition between organism and organism, between child and parent, will render this a very rare event; for in all cases the new and improved forms of life will tend to supplant the old and unimproved.

By the theory of natural selection all living species have been connected with the parent-species of each genus, by differences not greater than we see between the varieties of the same species at the present day; and these parent-species, now generally extinct, have in their turn been similarly connected with more ancient species; and so on backwards, always converging to the common ancestor of each great class.

So that the number of intermediate and transitional links, between all living and extinct species, must have been inconceivably great.

But assuredly, if this theory be true, such have lived upon this earth.