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Showing The Origin of Species Illustrated (The Illustrated Origin of Species) where title = '04-04 - Its Power at All Ages and on Both Sexes' order by subject, title, ordinal (2 Rows).
# subject title description
1
04 - Natural Selection
04-04 - Its Power at All Ages and on Both Sexes
As we see that those variations which, under domestication, appear at any particular period of life, tend to reappear in the offspring at the same period;- for instance, in the shape, size, and flavour of the seeds of the many varieties of our culinary and agricultural plants; in the caterpillar and cocoon stages of the varieties of the silk-worm; in the eggs of poultry, and in the colour of the down of their chickens; in the horns of our sheep and cattle when nearly adult;- so in a state of nature natural selection will be enabled to act on and modify organic beings at any age, by the accumulation of variations profitable at that age, and by their inheritance at a corresponding age.
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silkworm
silkworm

egg
egg

poultry
poultry

chicken
chicken

sheep
sheep

cattle
cattle


If it profit a plant to have its seeds more and more widely disseminated by the wind, I can see no greater difficulty in this being effected through natural selection, than in the cotton-planter increasing and improving by selection the down in the pods on his cotton-trees.

cotton
cotton


Natural selection may modify and adapt the larva of an insect to a score of contingencies, wholly different from those which concern the mature insect; and these modifications may affect, through correlation, the structure of the adult.

larva
larva

insect
insect


So, conversely, modifications in the adult may affect the structure of the larva; but in all cases natural selection will ensure that they shall not be injurious: for if they were so, the species would become extinct.
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2
04 - Natural Selection
04-04 - Its Power at All Ages and on Both Sexes
Natural selection will modify the structure of the young in relation to the parent, and of the parent in relation to the young.

In social animals it will adapt the structure of each individual for the benefit of the whole community, if the community profits by the selected change.

What natural selection cannot do, is to modify the structure of one species, without giving it any advantage, for the good of another species; and though statements to this effect may be found in works of natural history, I cannot find one case which will bear investigation.

A structure used only once in an animal's life, if of high importance to it, might be modified to any extent by natural selection; for instance, the great jaws possessed by certain insects, used exclusively for opening the cocoon- or the hard tip to the beak of unhatched birds, used for breaking the egg.

insect
insect

bird
bird

beak
beak

egg
egg


It has been asserted, that of the best short-beaked tumbler-pigeons a greater number perish in the egg than are able to get out of it; so that fanciers assist in the act of hatching.

Tumbler Pigeon
Tumbler Pigeon


Now if nature had to make the beak of a full-grown pigeon very short for the bird's own advantage, the process of modification would be very slow, and there would be simultaneously the most rigorous selection of all the young birds within the egg, which had the most powerful and hardest beaks, for all with weak beaks would inevitably perish; or, more delicate and more easily broken shells might be selected, the thickness of the shell being known to vary like every other structure.

It may be well here to remark that with all beings there must be much fortuitous destruction, which can have little or no influence on the course of natural selection.

For instance a vast number of eggs or seeds are annually devoured, and these could be modified through natural selection only if they varied in some manner which protected them from their enemies.

egg
egg

seeds
seeds


Yet many of these eggs or seeds would perhaps, if not destroyed, have yielded individuals better adapted to their conditions of life than any of these which happened to survive.

So again a vast number of mature animals and plants, whether or not they be the best adapted to their conditions, must be annually destroyed by accidental causes, which would not be in the least degree mitigated by certain changes of structure or constitution which would in other ways be beneficial to the species.

But let the destruction of the adults be ever so heavy, if the number which can exist in any district be not wholly kept down by such causes,- or again let the destruction of eggs or seeds be so great that only a hundredth or a thousandth part are developed,- yet of those which do survive, the best adapted individuals, supposing that there is any variability in favourable direction, will tend to propagate their kind in larger numbers than the less well adapted.

If the numbers be wholly kept down by the causes just indicated, as will often have been the case, natural selection will be powerless in certain beneficial directions; but this is no valid objection to its efficiency at other times and in other ways; for we are far from having any reason to suppose that many species ever undergo modification and improvement at the same time in the same area.
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Statistics and Drill Down Data Mining
subject #
01 - Variations Under Domestication 53 53
02 - Variations Under Nature 23 76
03 - Struggle for Existence 30 106
04 - Natural Selection 105 211
05 - Laws of Variation 48 259
06 - Difficutiles in Theory 74 333
07 - Instinct 13 346
08 - Hybridism 9 355
09 - On the Imperfection of the Geological Record 7 362
10 - On The Geological Succession of Organic Beings 10 372
11 - Geographical Distribution 7 379
12 - Geographical Distribution -- continued 23 402
13 - Mutual Affinities of Organic Beings: Morphology: Embryology: Rudimentary Or 34 436
14 - Recapitulation and Conclusion 38 474
title #
01-01 - Causes of Variability 5 5
01-02 - Effects of Habit 1 6
01-03 - correlation of Growth 2 8
01-04 - Inheritance 4 12
01-05 -Character of Domestic Varieties 2 14
01-06 - Difficulty of distinguishing between Varieties and Species 2 16
01-07 - Origin of Domestic Varieties from one or more Species 7 23
01-08 - Breeds of the Domestic Pigeons, their Differences and Origin 9 32
01-09 - Principles of Selection anciently followed, and their Effects 6 38
01-10 - Methodical and Unconscious Selection 5 43
01-11 - Unknown Origin of our Domestic Productions 5 48
01-12 - Circumstances favourable to Man's Power of Selection 3 51
01-13 - Summary 2 53
02-01 - Variability 2 55
02-02 - Individual Differences 2 57
02-03 - Doubtful Species 10 67
02-04 - Wide-ranging, much diffused, and common Species vary most 3 70
02-05 - Species of the Larger Genera in each Country vary more frequently than the Species of the Smaller Genera 2 72
02-06 - Many of the Species included within the Larger Genera resemble Varieties in being very closely, but unequally, related to each other, and in having restricted ranges 2 74
02-07 - Summary 2 76
03-01 - Bears on Natural Selection 2 78
03-02 - The Term, Struggle for Existence, used in a large sense 2 80
03-03 - Geometrical Ratio of Increase 2 82
03-04 - Rapid Increase of naturalised Animals and Plants 3 85
03-05 - Nature of the Checks to Increase 2 87
03-06 - Competition Universal 2 89
03-07 - Effects of Climate 2 91
03-08 - Protection from the Number of Individuals 2 93
03-09 - Complex Relations of all Animals and Plants Throughout Nature 5 98
03-10 - Struggle for Life most severe between Individuals and Varieties of the same Species 2 100
03-11 - The Relation of Organism to Organism the Most Important of All Relations 4 104
03-12 - Summary 2 106
04-01 - Natural Selection 5 111
04-02 - Its Power Compared with Man's Selection 2 113
04-03 - Its Power on Characters of Trifling Importance 2 115
04-04 - Its Power at All Ages and on Both Sexes 2 117
04-05 - Sexual Selection 3 120
04-06 - On the generality of Intercross Between Individuals of the Same Species 9 129
04-07 - Illustrations of the Action of Natural Selection: 10 139
04-08 - On the Intercrossing of Individuals 8 147
04-09 - Circumstances favourable for the production of new forms through Natural Selection 12 159
04-10 - Extinction caused by Natural Selection 3 162
04-11 - Divergence of Character 26 188
04-12 - On the Degree to which Organisation tends to advance 11 199
04-13 - Convergence of Character 8 207
04-14 - Summary of Chapter 4 211
05-01 - Effects of External Conditions 2 213
05-02 - Use and Disuse of Parts, combined with Natural Selection, Organs of Flight and Vision 7 220
05-03 - Acclimatisation 4 224
05-04 - Correlation of Growth 5 229
05-05 - Compensation and Economy of Growth 2 231
05-06 - False Correlation 2 233
05-07 - Multiple, Rudimentary, and Lowly-organised Structures are Variable 2 235
05-08 - Parts Developed in an Unusual Manner are Highly Variable 5 240
05-09 - Specific Characters more Variable than Generic Characters 2 242
05-10 - Secondary Sexual Characters Variable 3 245
05-11 - Species of the Same Genus Vary in an Analogous Manner 2 247
05-12 - Reversion to Long Lost Characters 10 257
05-13 - Summary 2 259
06-01 - Difficulties on the Theory of Descent with Modification 5 264
06-02 - Transitions 2 266
06-03 - Absence or Rarity of Transitional Varieties 10 276
06-04 - Transitions in Habits of Life 7 283
06-05 - Diversified Habits in the Same Species 2 285
06-06 - Species with Habits Widely Diffferent from those of their Allies 3 288
06-07 - Organs of extreme Perfection 5 293
06-08 - Means of Transition 6 299
06-09 - Cases of Difficulty 5 304
06-10 - Natura Non Facit Saltum 2 306
06-11 - Organs of Small Importance 6 312
06-12 - Organs not in all Cases Absolutely Perfect 13 325
06-13 - Summary: The Law of Unity of Type and of the Conditions of Existence Embraced by the Theory of Natural Selection 8 333
07-01 - Instincts comparable with habits, but different in their origin 2 335
07-02 - Instincts Graduated 2 337
07-03 - Aphides and ants 1 338
07-04 - Instincts variable 1 339
07-05 - Domestic instincts, their origin 1 340
07-06 - Natural instincts of the cuckoo, ostrich, and parasitic bees 1 341
07-07 - Slave-making ants 1 342
07-08 - Hive-bee, its cell-making instinct 1 343
07-09 - Difficulties on the theory of the Natural Selection of instincts 1 344
07-10 - Neuter or sterile insects 1 345
07-11 - Summary 1 346
08-01 - Distinction between the sterility of first crosses and of hybrids 1 347
08-02 - Sterility various in degree, not universal, affected by close interbreeding, removed by domestication 1 348
08-03 - Laws governing the sterility of hybrids 1 349
08-04 - Sterility not a special endowment, but incidental on other differences 1 350
08-05 - Causes of the sterility of first crosses and of hybrids 1 351
08-06 - Parallelism between the effects of changed conditions of life and crossing 1 352
08-07 - Fertility of varieties when crossed and of their mongrel offspring not universal 1 353
08-08 - Hybrids and mongrels compared independently of their fertility 1 354
08-09 - Summary 1 355
09-01 -On the absence of intermediate varieties at the present day 1 356
09-02 - On the nature of extinct intermediate varieties; on their number 1 357
09-03 - On the vast lapse of time, as inferred from the rate of deposition and of denudation 1 358
09-04 - On the poorness of our palaeontological collections 1 359
09-05 - On the intermittence of geological formations 1 360
09-06 - On the absence of intermediate varieties in any one formation 1 361
09-07 - On their sudden appearance in the lowest known fossiliferous strata 1 362
10-01 - On the slow and successive appearance of new species 1 363
10-02 - On their different rates of change 1 364
10-03 - Species once lost do not reappear 1 365
10-04 - Groups of species follow the same general rules in their appearance and disappearance as do single species 1 366
10-05 - On Extinction 1 367
10-06 - On simultaneous changes in the forms of life throughout the world 1 368
10-07 - On the affinities of extinct species to each other and to living species 1 369
10-08 - On the state of development of ancient forms 1 370
10-09 - On the succession of the same types within the same areas 1 371
10-10 - Summary of preceding and present chapters 1 372
11-01 - Present distribution cannot be accounted for by differences in physical conditions 1 373
11-02 - Importance of barriers 1 374
11-03 - Affinity of the productions of the same continent 1 375
11-04 - Centres of creation 1 376
11-05 - Means of dispersal, by changes of climate and of the level of the land, and by occasional means 2 378
11-06 - Dispersal during the Glacial period co-extensive with the world 1 379
12-10 - Distribution of fresh-water productions 1 380
12-20 - On the inhabitants of oceanic islands 1 381
12-30 - Absence of Batrachians and of terrestrial Mammals 4 385
12-40 - On the relations of the inhabitants of islands to those of the nearest mainland 3 388
12-50 - On colonisation from the nearest source with subsequent modification 10 398
12-60 - Summary of the last and present chapters 4 402
13-01 - CLASSIFICATION, groups subordinate to groups 1 403
13-02 - Natural system 1 404
13-03 - Rules and difficulties in classification, explained on the theory of descent with modification 13 417
13-04 - Classification of varieties 1 418
13-05 - Descent always used in classification 6 424
13-06 - Analogical or adaptive characters 2 426
13-07 - Affinities, general, complex and radiating 2 428
13-08 - Extinction separates and defines groups 4 432
13-09 - MORPHOLOGY, between members of the same class, between parts of the same individual 1 433
13-10 - EMBRYOLOGY, laws of, explained by variations not supervening at an early age, and being inherited at a corresponding age 1 434
13-11 - RUDIMENTARY ORGANS; their origin explained 1 435
13-12 - Summary 1 436
14-01 - Recapitulation of the difficulties on the theory of Natural Selection 7 443
14-02 - Recapitulation of the general and special circumstances in its favour 10 453
14-03 - Causes of the general belief in the immutability of species 13 466
14-04 - How far the theory of natural selection may be extended 1 467
14-05 - Effects of its adoption on the study of Natural history 5 472
14-06 - Concluding remarks 2 474
wolfs 1 475
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