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Showing The Origin of Species Illustrated (The Illustrated Origin of Species) where title = '04-05 - Sexual Selection' order by subject, title, ordinal (3 Rows).
# subject title description
1
04 - Natural Selection
04-05 - Sexual Selection
Inasmuch as peculiarities often appear under domestication in one sex and become hereditarily attached to that sex, so no doubt it will be under nature.

Thus it is rendered possible for the two sexes to be modified through natural selection in relation to different habits of life, as is sometimes the case; or for one sex to be modified in relation to the other sex, as commonly occurs.

This leads me to say a few words on what I have called Sexual Selection. This form of selection depends, not on a struggle for existence in relation to other organic beings or to external conditions, but on a struggle between the individuals of one sex, generally the males, for the possession of the other sex.

The result is not death to the unsuccessful competitor, but few or no offspring. Sexual selection is, therefore, less rigorous than natural selection.

Generally, the most vigorous males, those which are best fitted for their places in nature, will leave most progeny.

But in many cases, victory depends not so much on general vigor, as on having special weapons, confined to the male sex.

A hornless stag or spurless cock would have a poor chance of leaving numerous offspring.

stag
stag

cock
cock

spurs
spurs


Sexual selection, by always allowing the victor to breed, might surely give indomitable courage, length to the spur, and strength to the wing to strike in the spurred leg, in nearly the same manner as does the brutal cockfighter by the careful selection of his best cocks.

cockfight
cockfight


How low in the scale of nature the law of battle descends, I know not; male alligators have been described as fighting, bellowing, and whirling round, like Indians in a war-dance, for the possession of the females; male salmons have been observed fighting all day long; male stagbeetles sometimes bear wounds from the huge mandibles of other males; the males of certain hymenopterous insects have been frequently seen by that inimitable observer M. Fabre, fighting for a particular female who sits by, an apparently unconcerned beholder of the struggle, and then retires with the conqueror.

alligator
alligator

crocodile
crocodile

Stag Beetle
Stag Beetle

Indian
Indian


The war is, perhaps, severest between the males of polygamous animals, and these seem oftenest provided with special weapons.

The males of carnivorous animals are already well armed; though to them and to others, special means of defence may be given through means of sexual selection, as the mane of the lion, and the hooked jaw to the male salmon; for the shield may be as important for victory, as the sword or spear.

mane
mane

salmon
salmon

sword
sword

spear
spear
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2
04 - Natural Selection
04-05 - Sexual Selection
Amongst birds, the contest is often of a more peaceful character. All those who have attended to the subject, believe that there is the severest rivalry between the males of many species to attract, by singing, the females.

bird
bird


The rock-thrush of Guiana, birds of paradise, and some others, congregate; and successive males display with the most elaborate care, and show off in the best manner, their gorgeous plumage; they likewise perform strange antics before the females, which, standing by as spectators, at last choose the most attractive partner.

Rock Thrush
Rock Thrush

Rock Thrush (female)
Rock Thrush (female)

Bird of Paradise
Bird of Paradise

Bird of Paradise (flower)
Bird of Paradise (flower)


Those who have closely attended to birds in confinement well know that they often take individual preferences and dislikes: thus Sir R. Heron has described how a pied peacock was eminently attractive to all his hen birds.

peacock
peacock


I cannot here enter on the necessary details; but if man can in a short time give beauty and an elegant carriage to his bantams, according to his standard of beauty, I can see no good reason to doubt that female birds, by selecting, during thousands of generations, the most melodious or beautiful males, according to their standard of beauty, might produce a marked effect.

Some well-known laws, with respect to the plumage of male and female birds, in comparison with the plumage of the young, can partly be explained through the action of sexual selection on variations occurring at different ages, and transmitted to the males alone or to both sexes at corresponding ages; but I have not space here to enter on this subject.
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3
04 - Natural Selection
04-05 - Sexual Selection
Thus it is, as I believe, that when the males and females of any animal have the same general habits of life, but differ in structure, colour, or ornament, such differences have been mainly caused by sexual selection: that is, by individual males having had, in successive generations, some slight advantage over other males, in their weapons, means of defence, or charms, which they have transmitted to their male offspring alone.

Yet, I would not wish to attribute all sexual differences to this agency: for we see in our domestic animals peculiarities arising and becoming attached to the male sex, which apparently have not been augmented through selection by man.

The tuft of hair on the breast of the wild turkey-cock cannot be of any use, and it is doubtful whether it can be ornamental in the eyes of the female bird; indeed, had the tuft appeared under domestication, it would have been called a monstrosity.

Turkey Cock
Turkey Cock
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Statistics and Drill Down Data Mining
subject #
01 - Variations Under Domestication 53 53
02 - Variations Under Nature 23 76
03 - Struggle for Existence 30 106
04 - Natural Selection 105 211
05 - Laws of Variation 48 259
06 - Difficutiles in Theory 74 333
07 - Instinct 13 346
08 - Hybridism 9 355
09 - On the Imperfection of the Geological Record 7 362
10 - On The Geological Succession of Organic Beings 10 372
11 - Geographical Distribution 7 379
12 - Geographical Distribution -- continued 23 402
13 - Mutual Affinities of Organic Beings: Morphology: Embryology: Rudimentary Or 34 436
14 - Recapitulation and Conclusion 38 474
title #
01-01 - Causes of Variability 5 5
01-02 - Effects of Habit 1 6
01-03 - correlation of Growth 2 8
01-04 - Inheritance 4 12
01-05 -Character of Domestic Varieties 2 14
01-06 - Difficulty of distinguishing between Varieties and Species 2 16
01-07 - Origin of Domestic Varieties from one or more Species 7 23
01-08 - Breeds of the Domestic Pigeons, their Differences and Origin 9 32
01-09 - Principles of Selection anciently followed, and their Effects 6 38
01-10 - Methodical and Unconscious Selection 5 43
01-11 - Unknown Origin of our Domestic Productions 5 48
01-12 - Circumstances favourable to Man's Power of Selection 3 51
01-13 - Summary 2 53
02-01 - Variability 2 55
02-02 - Individual Differences 2 57
02-03 - Doubtful Species 10 67
02-04 - Wide-ranging, much diffused, and common Species vary most 3 70
02-05 - Species of the Larger Genera in each Country vary more frequently than the Species of the Smaller Genera 2 72
02-06 - Many of the Species included within the Larger Genera resemble Varieties in being very closely, but unequally, related to each other, and in having restricted ranges 2 74
02-07 - Summary 2 76
03-01 - Bears on Natural Selection 2 78
03-02 - The Term, Struggle for Existence, used in a large sense 2 80
03-03 - Geometrical Ratio of Increase 2 82
03-04 - Rapid Increase of naturalised Animals and Plants 3 85
03-05 - Nature of the Checks to Increase 2 87
03-06 - Competition Universal 2 89
03-07 - Effects of Climate 2 91
03-08 - Protection from the Number of Individuals 2 93
03-09 - Complex Relations of all Animals and Plants Throughout Nature 5 98
03-10 - Struggle for Life most severe between Individuals and Varieties of the same Species 2 100
03-11 - The Relation of Organism to Organism the Most Important of All Relations 4 104
03-12 - Summary 2 106
04-01 - Natural Selection 5 111
04-02 - Its Power Compared with Man's Selection 2 113
04-03 - Its Power on Characters of Trifling Importance 2 115
04-04 - Its Power at All Ages and on Both Sexes 2 117
04-05 - Sexual Selection 3 120
04-06 - On the generality of Intercross Between Individuals of the Same Species 9 129
04-07 - Illustrations of the Action of Natural Selection: 10 139
04-08 - On the Intercrossing of Individuals 8 147
04-09 - Circumstances favourable for the production of new forms through Natural Selection 12 159
04-10 - Extinction caused by Natural Selection 3 162
04-11 - Divergence of Character 26 188
04-12 - On the Degree to which Organisation tends to advance 11 199
04-13 - Convergence of Character 8 207
04-14 - Summary of Chapter 4 211
05-01 - Effects of External Conditions 2 213
05-02 - Use and Disuse of Parts, combined with Natural Selection, Organs of Flight and Vision 7 220
05-03 - Acclimatisation 4 224
05-04 - Correlation of Growth 5 229
05-05 - Compensation and Economy of Growth 2 231
05-06 - False Correlation 2 233
05-07 - Multiple, Rudimentary, and Lowly-organised Structures are Variable 2 235
05-08 - Parts Developed in an Unusual Manner are Highly Variable 5 240
05-09 - Specific Characters more Variable than Generic Characters 2 242
05-10 - Secondary Sexual Characters Variable 3 245
05-11 - Species of the Same Genus Vary in an Analogous Manner 2 247
05-12 - Reversion to Long Lost Characters 10 257
05-13 - Summary 2 259
06-01 - Difficulties on the Theory of Descent with Modification 5 264
06-02 - Transitions 2 266
06-03 - Absence or Rarity of Transitional Varieties 10 276
06-04 - Transitions in Habits of Life 7 283
06-05 - Diversified Habits in the Same Species 2 285
06-06 - Species with Habits Widely Diffferent from those of their Allies 3 288
06-07 - Organs of extreme Perfection 5 293
06-08 - Means of Transition 6 299
06-09 - Cases of Difficulty 5 304
06-10 - Natura Non Facit Saltum 2 306
06-11 - Organs of Small Importance 6 312
06-12 - Organs not in all Cases Absolutely Perfect 13 325
06-13 - Summary: The Law of Unity of Type and of the Conditions of Existence Embraced by the Theory of Natural Selection 8 333
07-01 - Instincts comparable with habits, but different in their origin 2 335
07-02 - Instincts Graduated 2 337
07-03 - Aphides and ants 1 338
07-04 - Instincts variable 1 339
07-05 - Domestic instincts, their origin 1 340
07-06 - Natural instincts of the cuckoo, ostrich, and parasitic bees 1 341
07-07 - Slave-making ants 1 342
07-08 - Hive-bee, its cell-making instinct 1 343
07-09 - Difficulties on the theory of the Natural Selection of instincts 1 344
07-10 - Neuter or sterile insects 1 345
07-11 - Summary 1 346
08-01 - Distinction between the sterility of first crosses and of hybrids 1 347
08-02 - Sterility various in degree, not universal, affected by close interbreeding, removed by domestication 1 348
08-03 - Laws governing the sterility of hybrids 1 349
08-04 - Sterility not a special endowment, but incidental on other differences 1 350
08-05 - Causes of the sterility of first crosses and of hybrids 1 351
08-06 - Parallelism between the effects of changed conditions of life and crossing 1 352
08-07 - Fertility of varieties when crossed and of their mongrel offspring not universal 1 353
08-08 - Hybrids and mongrels compared independently of their fertility 1 354
08-09 - Summary 1 355
09-01 -On the absence of intermediate varieties at the present day 1 356
09-02 - On the nature of extinct intermediate varieties; on their number 1 357
09-03 - On the vast lapse of time, as inferred from the rate of deposition and of denudation 1 358
09-04 - On the poorness of our palaeontological collections 1 359
09-05 - On the intermittence of geological formations 1 360
09-06 - On the absence of intermediate varieties in any one formation 1 361
09-07 - On their sudden appearance in the lowest known fossiliferous strata 1 362
10-01 - On the slow and successive appearance of new species 1 363
10-02 - On their different rates of change 1 364
10-03 - Species once lost do not reappear 1 365
10-04 - Groups of species follow the same general rules in their appearance and disappearance as do single species 1 366
10-05 - On Extinction 1 367
10-06 - On simultaneous changes in the forms of life throughout the world 1 368
10-07 - On the affinities of extinct species to each other and to living species 1 369
10-08 - On the state of development of ancient forms 1 370
10-09 - On the succession of the same types within the same areas 1 371
10-10 - Summary of preceding and present chapters 1 372
11-01 - Present distribution cannot be accounted for by differences in physical conditions 1 373
11-02 - Importance of barriers 1 374
11-03 - Affinity of the productions of the same continent 1 375
11-04 - Centres of creation 1 376
11-05 - Means of dispersal, by changes of climate and of the level of the land, and by occasional means 2 378
11-06 - Dispersal during the Glacial period co-extensive with the world 1 379
12-10 - Distribution of fresh-water productions 1 380
12-20 - On the inhabitants of oceanic islands 1 381
12-30 - Absence of Batrachians and of terrestrial Mammals 4 385
12-40 - On the relations of the inhabitants of islands to those of the nearest mainland 3 388
12-50 - On colonisation from the nearest source with subsequent modification 10 398
12-60 - Summary of the last and present chapters 4 402
13-01 - CLASSIFICATION, groups subordinate to groups 1 403
13-02 - Natural system 1 404
13-03 - Rules and difficulties in classification, explained on the theory of descent with modification 13 417
13-04 - Classification of varieties 1 418
13-05 - Descent always used in classification 6 424
13-06 - Analogical or adaptive characters 2 426
13-07 - Affinities, general, complex and radiating 2 428
13-08 - Extinction separates and defines groups 4 432
13-09 - MORPHOLOGY, between members of the same class, between parts of the same individual 1 433
13-10 - EMBRYOLOGY, laws of, explained by variations not supervening at an early age, and being inherited at a corresponding age 1 434
13-11 - RUDIMENTARY ORGANS; their origin explained 1 435
13-12 - Summary 1 436
14-01 - Recapitulation of the difficulties on the theory of Natural Selection 7 443
14-02 - Recapitulation of the general and special circumstances in its favour 10 453
14-03 - Causes of the general belief in the immutability of species 13 466
14-04 - How far the theory of natural selection may be extended 1 467
14-05 - Effects of its adoption on the study of Natural history 5 472
14-06 - Concluding remarks 2 474
wolfs 1 475
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