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Showing The Origin of Species Illustrated (The Illustrated Origin of Species) where title = '04-11 - Divergence of Character' order by subject, title, ordinal limit 0, 4 (4 of 26).
# subject title description
1
04 - Natural Selection
04-11 - Divergence of Character
The principle, which I have designated by this term, is of high importance, and explains, as I believe, several important facts.

In the first place, varieties, even strongly-marked ones, though having somewhat of the character of species- as is shown by the hopeless doubts in many cases how to rank them- yet certainly differ far less from each other than do good and distinct species.

Nevertheless, according to my view, varieties are species in the process of formation, or are, as I have called them, incipient species.

How, then, does the lesser difference between varieties become augmented into the greater difference between species?

That this does habitually happen, we must infer from most of the innumerable species throughout nature presenting well-marked differences; whereas varieties, the supposed prototypes and parents of future well-marked species, present slight and ill-defined differences.

Mere chance, as we may call it, might cause one variety to differ in some character from its parents, and the offspring of this variety again to differ from its parent in the very same character and in a greater degree; but this alone would never account for so habitual and large a degree of difference as that between the species of the same genus.
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2
04 - Natural Selection
04-11 - Divergence of Character
As has always been my practice, I have sought light on this head from our domestic productions.

We shall here find something analogous.

It will be admitted that the production of races so different as short-horn and Hereford cattle, race and cart horses, the several breeds of pigeons, &c., could never have been effected by the mere chance accumulation of similar variations during many successive generations.

Hereford Cow
Hereford Cow

Race Horse
Race Horse


In practice, a fancier is, for instance, struck by a pigeon having a slightly shorter beak; another fancier is struck by a pigeon having a rather longer beak; and on the acknowledged principle that "fanciers do not and will not admire a medium standard, but like extremes," they both go on (as has actually occurred with the sub-breeds of the tumbler-pigeon) choosing and breeding from birds with longer and longer beaks, or with shorter and shorter beaks.

pigeon
pigeon


Again, we may suppose that at an early period of history, the men of one nation or district required swifter horses, whilst those of another required stronger and bulkier horses.

Dray Horse
Dray Horse


The early differences would be very slight; but, in the course of time from the continued selection of swifter horses in the one case, and of stronger ones in the other, the differences would become greater, and would be noted as forming two sub-breeds.

Ultimately, after the lapse of centuries, these sub-breeds would become converted into two well-established and distinct breeds.

As the differences became greater, the inferior animals with intermediate characters, being neither swift nor very strong, would not have been used for breeding, and will thus have tended to disappear.

Here, then, we see in man's productions the action of what may be called the principle of divergence, causing differences, at first barely appreciable, steadily to increase, and the breeds to diverge in character, both from each other and from their common parent.
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3
04 - Natural Selection
04-11 - Divergence of Character
But how, it may be asked, can any analogous principle apply in nature?

I believe it can and does apply most efficiently (though it was a long time before I saw how), from the simple circumstance that the more diversified the descendants from any one species become in structure, constitution, and habits, by so much will they be better enabled to seize on many and widely diversified places in the polity of nature, and so be enabled to increase in numbers.
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4
04 - Natural Selection
04-11 - Divergence of Character
We can clearly discern this in the case of animals with simple habits.

Take the case of a carnivorous quadruped, of which the number that can be supported in any country has long ago arrived at its full average.

If its natural power of increase be allowed to act, it can succeed in increasing (the country not undergoing any change in conditions) only by its varying descendants seizing on places at present occupied by other animals: some of them, for instance, being enabled to feed on new kinds of prey, either dead or alive; some inhabiting new stations, climbing trees, frequenting water, and some perhaps becoming less carnivorous.

quadruped
quadruped

carnivore
carnivore

herbivore
herbivore


The more diversified in habits and structure the descendants of our carnivorous animals become, the more places they will be enabled to occupy.

What applies to one animal will apply throughout all time to all animals- that is, if they vary- for otherwise natural selection can effect nothing.

So it will be with plants. It has been experimentally proved, that if a plot of ground be sown with one species of grass, and a similar plot be sown with several distinct genera of grasses, a greater number of plants and a greater weight of dry herbage can be raised in the latter than in the former case.

grass
grass


The same has been found to hold good when one variety and several mixed varieties of wheat have been sown on equal spaces of ground.

wheat
wheat


Hence, if any one species of grass were to go on varying, and the varieties were continually selected which differed from each other in the same manner, though in a very slight degree, as do the distinct species and genera of grasses, a greater number of individual plants of this species, including its modified descendants, would succeed in living on the same piece of ground.

And we know that each species and each variety of grass is annually sowing almost countless seeds; and is thus striving, as it may be said, to the utmost to increase in number.

Consequently, in the course of many thousand generations, the most distinct varieties of any one species of grass would have the best chance of succeeding and of increasing in numbers, and thus of supplanting the less distinct varieties; and varieties, when rendered very distinct from each other, take the rank of species.

grass
grass
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Page 1 of 7 (4 rows per page)
1 2 3 4 5 6 7

Statistics and Drill Down Data Mining
subject #
01 - Variations Under Domestication 22 22
02 - Variations Under Nature 23 45
03 - Struggle for Existence 30 75
04 - Natural Selection 104 179
05 - Laws of Variation 47 226
06 - Difficutiles in Theory 74 300
07 - Instinct 13 313
08 - Hybridism 8 321
09 - On the Imperfection of the Geological Record 7 328
10 - On The Geological Succession of Organic Beings 10 338
11 - Geographical Distribution 6 344
12 - Geographical Distribution -- continued 23 367
13 - Mutual Affinities of Organic Beings: Morphology: Embryology: Rudimentary Or 34 401
14 - Recapitulation and Conclusion 38 439
title #
01-08 - Breeds of the Domestic Pigeons, their Differences and Origin 1 1
01-09 - Principles of Selection anciently followed, and their Effects 6 7
01-10 - Methodical and Unconscious Selection 5 12
01-11 - Unknown Origin of our Domestic Productions 5 17
01-12 - Circumstances favourable to Man's Power of Selection 3 20
01-13 - Summary 2 22
02-01 - Variability 2 24
02-02 - Individual Differences 2 26
02-03 - Doubtful Species 10 36
02-04 - Wide-ranging, much diffused, and common Species vary most 3 39
02-05 - Species of the Larger Genera in each Country vary more frequently than the Species of the Smaller Genera 2 41
02-06 - Many of the Species included within the Larger Genera resemble Varieties in being very closely, but unequally, related to each other, and in having restricted ranges 2 43
02-07 - Summary 2 45
03-01 - Bears on Natural Selection 2 47
03-02 - The Term, Struggle for Existence, used in a large sense 2 49
03-03 - Geometrical Ratio of Increase 2 51
03-04 - Rapid Increase of naturalised Animals and Plants 3 54
03-05 - Nature of the Checks to Increase 2 56
03-06 - Competition Universal 2 58
03-07 - Effects of Climate 2 60
03-08 - Protection from the Number of Individuals 2 62
03-09 - Complex Relations of all Animals and Plants Throughout Nature 5 67
03-10 - Struggle for Life most severe between Individuals and Varieties of the same Species 2 69
03-11 - The Relation of Organism to Organism the Most Important of All Relations 4 73
03-12 - Summary 2 75
04-01 - Natural Selection 5 80
04-02 - Its Power Compared with Man's Selection 2 82
04-03 - Its Power on Characters of Trifling Importance 2 84
04-04 - Its Power at All Ages and on Both Sexes 2 86
04-05 - Sexual Selection 2 88
04-06 - On the generality of Intercross Between Individuals of the Same Species 9 97
04-07 - Illustrations of the Action of Natural Selection: 10 107
04-08 - On the Intercrossing of Individuals 8 115
04-09 - Circumstances favourable for the production of new forms through Natural Selection 12 127
04-10 - Extinction caused by Natural Selection 3 130
04-11 - Divergence of Character 26 156
04-12 - On the Degree to which Organisation tends to advance 11 167
04-13 - Convergence of Character 8 175
04-14 - Summary of Chapter 4 179
05-01 - Effects of External Conditions 2 181
05-02 - Use and Disuse of Parts, combined with Natural Selection, Organs of Flight and Vision 7 188
05-03 - Acclimatisation 4 192
05-04 - Correlation of Growth 5 197
05-05 - Compensation and Economy of Growth 2 199
05-06 - False Correlation 1 200
05-07 - Multiple, Rudimentary, and Lowly-organised Structures are Variable 2 202
05-08 - Parts Developed in an Unusual Manner are Highly Variable 5 207
05-09 - Specific Characters more Variable than Generic Characters 2 209
05-10 - Secondary Sexual Characters Variable 3 212
05-11 - Species of the Same Genus Vary in an Analogous Manner 2 214
05-12 - Reversion to Long Lost Characters 10 224
05-13 - Summary 2 226
06-01 - Difficulties on the Theory of Descent with Modification 5 231
06-02 - Transitions 2 233
06-03 - Absence or Rarity of Transitional Varieties 10 243
06-04 - Transitions in Habits of Life 7 250
06-05 - Diversified Habits in the Same Species 2 252
06-06 - Species with Habits Widely Diffferent from those of their Allies 3 255
06-07 - Organs of extreme Perfection 5 260
06-08 - Means of Transition 6 266
06-09 - Cases of Difficulty 5 271
06-10 - Natura Non Facit Saltum 2 273
06-11 - Organs of Small Importance 6 279
06-12 - Organs not in all Cases Absolutely Perfect 13 292
06-13 - Summary: The Law of Unity of Type and of the Conditions of Existence Embraced by the Theory of Natural Selection 8 300
07-01 - Instincts comparable with habits, but different in their origin 2 302
07-02 - Instincts Graduated 2 304
07-03 - Aphides and ants 1 305
07-04 - Instincts variable 1 306
07-05 - Domestic instincts, their origin 1 307
07-06 - Natural instincts of the cuckoo, ostrich, and parasitic bees 1 308
07-07 - Slave-making ants 1 309
07-08 - Hive-bee, its cell-making instinct 1 310
07-09 - Difficulties on the theory of the Natural Selection of instincts 1 311
07-10 - Neuter or sterile insects 1 312
07-11 - Summary 1 313
08-01 - Distinction between the sterility of first crosses and of hybrids 1 314
08-03 - Laws governing the sterility of hybrids 1 315
08-04 - Sterility not a special endowment, but incidental on other differences 1 316
08-05 - Causes of the sterility of first crosses and of hybrids 1 317
08-06 - Parallelism between the effects of changed conditions of life and crossing 1 318
08-07 - Fertility of varieties when crossed and of their mongrel offspring not universal 1 319
08-08 - Hybrids and mongrels compared independently of their fertility 1 320
08-09 - Summary 1 321
09-01 -On the absence of intermediate varieties at the present day 1 322
09-02 - On the nature of extinct intermediate varieties; on their number 1 323
09-03 - On the vast lapse of time, as inferred from the rate of deposition and of denudation 1 324
09-04 - On the poorness of our palaeontological collections 1 325
09-05 - On the intermittence of geological formations 1 326
09-06 - On the absence of intermediate varieties in any one formation 1 327
09-07 - On their sudden appearance in the lowest known fossiliferous strata 1 328
10-01 - On the slow and successive appearance of new species 1 329
10-02 - On their different rates of change 1 330
10-03 - Species once lost do not reappear 1 331
10-04 - Groups of species follow the same general rules in their appearance and disappearance as do single species 1 332
10-05 - On Extinction 1 333
10-06 - On simultaneous changes in the forms of life throughout the world 1 334
10-07 - On the affinities of extinct species to each other and to living species 1 335
10-08 - On the state of development of ancient forms 1 336
10-09 - On the succession of the same types within the same areas 1 337
10-10 - Summary of preceding and present chapters 1 338
11-01 - Present distribution cannot be accounted for by differences in physical conditions 1 339
11-02 - Importance of barriers 1 340
11-03 - Affinity of the productions of the same continent 1 341
11-04 - Centres of creation 1 342
11-05 - Means of dispersal, by changes of climate and of the level of the land, and by occasional means 1 343
11-06 - Dispersal during the Glacial period co-extensive with the world 1 344
12-10 - Distribution of fresh-water productions 1 345
12-20 - On the inhabitants of oceanic islands 1 346
12-30 - Absence of Batrachians and of terrestrial Mammals 4 350
12-40 - On the relations of the inhabitants of islands to those of the nearest mainland 2 352
12-50 - On colonisation from the nearest source with subsequent modification 11 363
12-60 - Summary of the last and present chapters 4 367
13-01 - CLASSIFICATION, groups subordinate to groups 1 368
13-02 - Natural system 1 369
13-03 - Rules and difficulties in classification, explained on the theory of descent with modification 13 382
13-04 - Classification of varieties 1 383
13-05 - Descent always used in classification 6 389
13-06 - Analogical or adaptive characters 2 391
13-07 - Affinities, general, complex and radiating 2 393
13-08 - Extinction separates and defines groups 4 397
13-09 - MORPHOLOGY, between members of the same class, between parts of the same individual 1 398
13-10 - EMBRYOLOGY, laws of, explained by variations not supervening at an early age, and being inherited at a corresponding age 1 399
13-11 - RUDIMENTARY ORGANS; their origin explained 1 400
13-12 - Summary 1 401
14-01 - Recapitulation of the difficulties on the theory of Natural Selection 7 408
14-02 - Recapitulation of the general and special circumstances in its favour 10 418
14-03 - Causes of the general belief in the immutability of species 13 431
14-04 - How far the theory of natural selection may be extended 1 432
14-05 - Effects of its adoption on the study of Natural history 5 437
14-06 - Concluding remarks 2 439
wolfs 1 440
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